Fennel fronds share enough flavor and texture with a few other feathery herbs that substitution works well in both directions - useful whether you're out of fronds and need a replacement, or have fronds on hand and want to use them in place of something else.
Dill is the closest substitute for fennel fronds in both appearance and general character - feathery, mild, and slightly sweet, though without fennel's specific anise note. Use a roughly equal amount in most recipes, including our yogurt sauce or frond oil, where the swap works particularly well.
Tarragon carries a more pronounced anise flavor, closer in spirit to fennel than dill is, but it's also considerably stronger, so use about half the amount called for and adjust from there. It works especially well as a substitute in fennel frond pesto, where a bolder anise character isn't a problem.
If you have neither fresh fronds nor a good substitute herb, a small amount of crushed fennel seed (used sparingly, since it's far more concentrated) can approximate some of the same flavor in a cooked dish, though it won't replicate the fresh, leafy texture fronds bring to something like a salad or garnish.
Fennel fronds work well as a substitute anywhere fresh dill is called for, particularly in yogurt-based sauces, fish dishes, and pickling brines. The flavor isn't identical, but it's close enough that most dill-forward recipes hold up well with the swap.
Fronds are considerably milder than tarragon, so a straight swap will result in a noticeably less intense anise flavor - use roughly double the amount of fronds to approximate tarragon's presence in a dish, and taste as you go rather than measuring precisely.
This works best in pesto specifically, as in our fennel frond pesto recipe, where the general format of blended herb, nuts, cheese, and oil holds up regardless of which specific herb anchors the sauce. It's a less natural fit as a direct basil substitute in most other basil-forward dishes, like a caprese salad, where basil's specific peppery character is more central to the dish.
Knowing these substitutions both ways makes fennel fronds, dill, and tarragon feel less like three separate specialty herbs and more like a flexible family you can move between depending on what's actually in the fridge. For more on fennel fronds specifically, see the complete fennel fronds guide.